A study on COM motion in three fish species with different morphologies discovered that the surge fluctuated at twice frequency of the tail for eels and sunfish, while the sway resulted at the same frequency as the tail beat. Despite surge oscillations not changing significantly with increasing speed, sway oscillations did increase. Other study using particle image velocimetry discovered that the water flow of fish with homocercal tails produces vortex rings in a linked-chain pattern, while fish with heterocercal tails like sharks produce dual linked-ring vortexes. Regarding the effect of skin denticles on performance, a study discovered that intact shark skin provides 12.3% more speed than membranes with removed denticles. Lastly, a study using robotic models found that small length changes in swimming bodies can alter speed by a factor of three to five times.
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